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Chapter 33
Significant Ecological Resources and Endangered Species
33.1 Ecological Significance of the Cape Fear River Basin
The Cape Fear River basin is the largest of North Carolina’s river basins, and because of its size
it contains a wide variety of aquatic systems. The Cape Fear River itself has the character of
three or more rivers including: the clearwater Piedmont stream that rises at the confluence of the
Deep River and the Haw River; a meandering coastal “brownwater” river draining farmlands at
its mid-section; and a 30-mile long brackish estuary at its lower end. Also in the basin are
“blackwater” tributaries such as the Black, South and Northeast Cape Fear Rivers. Not only is
the Cape Fear River basin high in natural diversity, it also has a high rate of endemism. Among
the many rare mussels and fish known from the basin are species found nowhere else.
33.2 Rare Aquatic and Wetland-Dwelling Animal Species
For information on any of the species listed in Table 35, visit the NC Natural Heritage Program
website at www.ncsparks.net/nhp.
Table 35 List of Rare Species associated with Aquatic Habitats in the Cape Fear River
Basin
Group Scientific
Name
Common
Name
State
Status
Federal
Status
Crustacean Cambarus catagius Greensboro burrowing crayfish SC
Crustacean Cambarus davidi Carolina ladle crayfish SR
Fish Evorthodus lyricus Lyre goby SR
Fish Acipenser brevirostrum Shortnose sturgeon E E
Fish Fundulus luciae Spotfin killifish SR
Fish Etheostoma collis pop 2 Carolina darter - eastern piedmont population SC FSC
Fish Carpiodes velifer Highfin carpsucker SC
Fish Lucania goodei Bluefin killifish SC
Fish Heterandria formosa Least killifish SC
Fish Notropis mekistocholas Cape Fear shiner E E
Fish Hypsoblennius ionthas Freckled blenny SR
Fish Microphis brachyurus Opossum pipefish SR
Fish Poecilia latipinna Sailfin molly SR
Fish Noturus sp. 1 Broadtail madtom SC
Fish Eleotris pisonis Spinycheek sleeper SR
Chapter 33 – Significant Ecological Resources and Endangered Species 298
Fish Elassoma boehlkei Carolina pygmy sunfish T FSC
Fish Cyprinella zanema pop 2 Santee chub - Coastal Plain population SC
Fish Semotilus lumbee Sandhills chub SC
Gobionellus stigmaticus SR
Moxostoma sp. 2 SR FSC
Fish Ambloplites cavifrons Roanoke bass SR
Insect Ephemerella argo Argo ephemerellan mayfly SR FSC
Insect Tricorythodes robacki A mayfly SR
Insect Gomphus septima Septima's clubtail SR FSC
Insect Triaenodes marginata A triaenode caddisfly SR
Insect Dolania americana American sand burrowing mayfly SR FSC
Insect Progomphus bellei Belle's sanddragon SR FSC
Insect Choroterpes basalis A mayfly SR
Insect Ceraclea cancellata A caddisfly SR
Mollusk Villosa vaughaniana Carolina creekshell E FSC
Mollusk Fusconaia masoni Atlantic pigtoe FSC E
Magnificent rams-horn E FSC
Anodonta couperiana Barrel floater E
Reptile Alligator mississippiensis American alligator T T(S/A)
Reptile Caretta caretta Loggerhead seaturtle T T
Reptile Malaclemys terrapin
centrata Carolina diamond-backed terrapin SC
Fish Marked goby
Fish Carolina redhorse
Mollusk Planorbella magnifica
Mollusk
Rare Species Listing Criteria
E = Endangered (those species in danger of becoming extinct)
T = Threatened (considered likely to become endangered within the foreseeable future)
SR = Significantly Rare (those whose numbers are small and whose populations need monitoring)
SC = Species of Special Concern
FSC = Federal Species of Concern (those under consideration for listing under the Federal Endangered Species Act)
T(S/A) = Threatened due to similarity of appearance
EX = Extirpated
33.3 Significant Natural Heritage Areas in the Cape Fear River Basin
Figure 42 is a map of the Significant Natural Heritage Areas of the Cape Fear River basin. The
Natural Heritage Program identifies sites (terrestrial or aquatic) that have particular biodiversity
significance. A site’s significance may be due to the presence of rare species, rare or high
quality natural communities, or other important ecological features. The accompanying map
shows the Significant Natural Heritage Areas identified in the Cape Fear River basin. Over 450
individual natural areas have been identified in the Cape Fear River basin, too large a number to
discuss in detail here, so only the most important of these areas are discussed below.
Chapter 33 – Significant Ecological Resources and Endangered Species 299
33.3.1 Cape Fear Shiner Aquatic Habitats
Sections of three Piedmont rivers form the primary population centers for the very rare and
endangered Cape Fear shiner (Notropis mekistocholas), a fish endemic to a small part of the
Cape Fear River basin. Because of this, stretches of these rivers have been designated by the US
Fish and Wildlife Service as critical habitat. These rivers, the Deep, Rocky and Haw, also
support a number of other rare aquatic animals, including fish such as the Carolina redhorse
(Moxostoma sp.) and the Roanoke bass (Ambloplites cavifrons); freshwater mussels such as the
brook floater (Alasmidonta varicosa), triangle floater (Alasmidonta undulata), Atlantic pigtoe
(Fusconaia masoni), yellow lampmussel (Lampsilis cariosa), squawfoot (Strophitus undulatus),
notched rainbow (Villosa constricta), and eastern creekshell (Villosa delumbis); and dragonflies
such as Septima's clubtail (Gomphus septima). Also found here is one of just two North Carolina
populations of the endangered plant harperella (Ptilimnium nodosum), found on an island in the
Deep River in Chatham County.
33.3.2 Sandhills Megasite
The diverse natural communities of the sandhills region, such as hillside seeps, upland longleaf
pine forests, streamhead pocosins, and mixed hardwood-Atlantic white cedar swamps, provide
habitat for many rare and endemic species including perhaps the largest remaining concentration
of the endangered red-cockaded woodpecker (Picoides borealis) in the state. Endangered plants
include Michaux's sumac (Rhus michauxii), chaffseed (Schwalbea americana), and rough-leaf
loosestrife (Lysimachia asperulifolia). The publicly-owned Sandhills Game Land, Camp
MacKall and Fort Bragg contain some of the largest remnants of the sandhills in natural
condition in North Carolina.
33.3.3 Bladen Lakes Megasite
Occurring on ancient terraces of the Cape Fear River, the Bladen Lakes area contains the greatest
concentration of relatively unaltered Carolina bays in North Carolina. The bays contain lakes
and a diversity of peatland communities. The surrounding landscape is an irregular mosaic of
shallow peatlands and sand longleaf pine communities. Many of the significant natural areas are
in public ownership, either as game lands, state forest or state parks.
33.3.4 Black and South Rivers
The Black and South Rivers contain significant aquatic communities with two rare fish species –
the broadtail madtom (Noturus sp.) and the Santee chub (Cyprinella zanema); and several rare
mussels – pod lance (Elliptio folliculata), Cape Fear spike (E. marsupiobesa), Atlantic pigtoe
(Fusconaia masoni), and yellow lampmussel (Lampsilis cariosa). The Black River is also one of
the best remaining examples of a blackwater river system in the southeast coastal plain.
Particularly notable is an ancient cypress-gum swamp, which contains the oldest stand of trees
east of the Rocky Mountains; some cypress trees in this swamp have been core-dated to 364 AD.
The swamp forest of the Black River supports several colonies of the rare Rafinesque's big-eared
bat (Corynorhinus rafinesquii), a bat that is dependent on large hollow trees found in old-growth
forests.
Chapter 33 – Significant Ecological Resources and Endangered Species 300
33.3.5 Lower Cape Fear River
The lower reach of the Cape Fear River is brackish and supports numerous rare marine fishes,
including the endangered shortnose sturgeon (Acipenser brevirostrum), as well as freckled
blennies (Hypsoblennius ionthas), marked gobies (Gobionellus stigmaticus), spinycheek sleepers
(Eleotris pisonis), and opossum pipefish (Microphis brachyurus). The endangered manatee
(Trichechus manatus) is an occasional visitor, especially in summer.
33.3.6 Northeast Cape Fear River Floodplain
The Northeast Cape Fear River floodplain is a drowned blackwater river corridor characterized
by Tidal Freshwater Marshes at the lower end and extensive Tidal Cypress--Gum Swamp
communities upstream. Encompassing more than 22,000 acres, the Northeast Cape Fear River
floodplain contains some very high quality natural communities, including rare types such as
Peatland Atlantic White Cedar Forest and longleaf pine forests. At least one portion of the site
contains old-growth longleaf pine communities and mature examples of nonriverine wetland
communities. Though not thoroughly explored, a number of rare plant and animal species have
been found here. The southeastern myotis (Myotis austroriparius) – a bat that roosts in hollow
trees near water – is widespread along the upper portion, and alligators are present throughout.
The rare estuarine fishes lyre goby (Evorthodus lyricus) and spinycheek sleeper (Eleotris
pisonis) have been found in the marshes at the lower end. The site is a major forested connector
between other large sites.
33.3.7 Pine Savannas
The Cape Fear River basin contains a high concentration of savanna habitats. These wet,
generally flat areas have an open to sparse tree canopy, with few shrubs and a dense herb layer.
The pine savannas of southeastern North Carolina have among the highest species diversity of
any natural community in temperate North America, and more rare species are associated with
the pine savanna than any other natural community type in the state. Only a few examples of
this natural community are protected through public ownership or conservation easement.
33.3.8 Holly Shelter/Angola Bay Megasite
The Holly Shelter/Angola Bay region is one of the largest, nearly contiguous natural areas in the
state. The Holly Shelter Game Land has a large domed peatland in its center that supports one of
the largest pocosin community complexes in the state. On the southeast side of the Game Land
is an extensive relict beach ridge system with associated longleaf pine communities and great
concentrations of rare species, including one of the largest populations of Venus flytrap
(Dionaea muscipula) in the state and several dozen red-cockaded woodpecker colonies. To the
north is Angola Bay, one of states most inaccessible interior wetlands. The bay has a diameter of
more than ten miles with no roads or trails. Most of the area is low pocosin with scattered pond
pine in an otherwise near-treeless expanse of evergreen shrubbery.
Chapter 33 – Significant Ecological Resources and Endangered Species 301
33.3.9 Boiling Spring Lakes Wetland Complex
The Boiling Spring Lakes Wetland Complex is another large, hydrologically intact complex of
relict beach ridges and swales interspersed with Carolina bays. The extensive pine flatwoods,
pine savannas, pond pine woodlands, and pocosin communities of this area support a number of
rare plant species. This ecologically significant natural area was until recently the largest
unprotected natural longleaf pine/pocosin landscape complex in the state. However, the NC
Department of Agriculture and The Nature Conservancy are currently focusing protection efforts
in this area.
33.4 Significant Aquatic Habitats in Cape Fear River Basin
The Natural Heritage Program collaborates with a number of freshwater ecologists in other
agencies and organizations to identify Significant Aquatic Habitats in North Carolina.
Significant Aquatic Habitats are stream segments or other bodies of water that contain significant
natural resources, such as a high diversity of rare aquatic animal species. The impact from lands
adjacent and upstream of these stream reaches will determine their water quality and the viability
of their aquatic species. The Significant Aquatic Habitats of the Cape Fear River basin include:
Bear Creek Aquatic Habitat - A regionally-significant creek that adjoins the Rocky River-Lower
Deep River Aquatic Habitat at its downstream end. This medium-sized stream supports a
diverse mussel fauna, including the brook floater (Alasmidonta varicosa) and Atlantic elktoe
(Fusconaia masoni).
Black River Aquatic Habitat - A state-significant aquatic habitat containing four rare mussels:
Cape Fear Spike (Elliptio marsupiobesa), Atlantic pigtoe (Fusconaia masoni), yellow
lampmussel (Lampsilis cariosa), and pod lance (Elliptio folliculata). Other rare aquatic animals
known from the Black River include river frog (Rana hecksheri), American alligator, and the
American sand burrowing mayfly (Dolania americana). The American sand burrowing mayfly
is a predaceous mayfly that is able to dig rapidly into sandy substrates. It is known from only a
few places in the southeast, and in North Carolina has only been found in one location – the
Black River.
Deep River (Moore/Randolph) Aquatic Habitat - Considered to be nationally significant, it is
home to a population of the Cape Fear Shiner (Notropis mekistocholas), as well as the Carolina
redhorse (Moxostoma sp. 2), Roanoke bass (Ambloplites cavifrons), and a number of mussels,
including triangle floater (Alasmidonta undulata), brook floater (Alasmidonta varicosa), Atlantic
pigtoe (Fusconia masoni), yellow lampmussel (Lampsilis cariosa), squawfoot (Strophitus
undulatus), and notched rainbow (Villosa constricta).
Haw River Aquatic Habitat - The nationally significant stretch of the Haw River contains one of
just a few known sites for the endemic and Federally Endangered Cape Fear shiner (Notropis
mekistocholas). The globally rare Septima's clubtail (Gomphus septima) is also found here.
Lower Cape Fear River Aquatic Habitat - The state significant lower Cape Fear River is brackish
and contains numerous rare animals. Records of the shortnose sturgeon indicate that it occurs
here rarely, while manatees are found more occasionally, especially in summer. Alligators are
Chapter 33 – Significant Ecological Resources and Endangered Species 303
present mainly in tributary streams. Freckled blenny and spinycheek sleeper are rare marine
fishes of this section of the river.
Rocky River/Lower Deep River Aquatic Habitat - This nationally-significant habitat is separated
from the Upper Rocky River Aquatic Habitat (see below) by an impoundment. The federally
endangered Cape Fear (Notropis mekistocholas) shiner has its primary population center in this
area. The aquatic plant harperella (Ptilimnium nodosum) only has two known populations in
North Carolina, one of which occurs on an island in the Deep River.
South River Aquatic Habitat - A state-significant site discussed above in association with the
Black River. This high quality blackwater river contains two undescribed and rare fish species,
Noturus sp. and Hybopsis sp., and diversity of other aquatic biota.
Town Creek Aquatic Habitat - This nationally-significant site is a short creek that flows eastward
in eastern Brunswick County and empties into the Cape Fear River. Despite its short length, it
contains the only known population of the Greenfield ramshorn snail (Helisoma eucosmium), a
globally rare and imperiled mollusk, as well as several other rare animals and plants.
University Lake Aquatic Habitat - This state-significant site contains the largest population in the
state, by far, of the rare mussel Savannah lilliput (Toxolasma pullus).
Upper Cape Fear River Aquatic Habitat - This state-significant site passes through Harnett,
Chatham and Lee counties. This site contains seven rare mussels and two rare fishes: the
Carolina redhorse (Moxostoma sp. 2) and occasional reports of the federally endangered Cape
Fear shiner (Notropis mekistocholas).
Upper Rocky River Aquatic Habitat - This state-significant site is separated from the Rocky
River/Lower Deep River Aquatic Habitat (see above) by the Reeves Lake impoundment. The
site has a population of the federally endangered Cape Fear shiner (Notropis mekistocholas),
along with two rare mollusks and one rare stream insect.
Upper Black River Aquatic Habitat - This site contains lower portions of Six Runs and Great
Coharie and Little Coharie Creeks, where biologists have found three rare mussels: eastern
creekshell (Villosa delumbis), eastern lampmussel (Lampsilis radiata radiata), and pod lance
(Elliptio folliculata), as well as two rare fishes: Santee chub (Cyprinella zanema) and broadtail
madtom (Noturus sp. 1).
There are a number of Significant Natural Heritage Areas not listed here that contribute to Cape
Fear River water quality. Please contact the NC Natural Heritage Program to obtain information
about these natural areas. Significant Natural Heritage Areas are identified by the Natural
Heritage Program, but the identification of a natural area conveys no protection. Protection
comes from the landowner. For details about any of the Significant Natural Heritage Areas,
please contact the North Carolina Natural Heritage Program.
Chapter 33 – Significant Ecological Resources and Endangered Species 304
33.5 Public Lands
The accompanying map shows the land protected by public ownership in the Cape Fear River
basin. Many significant natural areas, including some already mentioned, are located on public
land. Also on the map are some preserves or conservation easements held by private
conservation organizations.
Some notable public lands within the Cape Fear River basin include the military lands – Fort
Bragg, Pope Air Force Base, Military Ocean Terminal Sunny Point, and part of Camp Lejeune –
substantial areas which balance conservation with military training missions. Another large
piece of land in the Cape Fear River basin is Jordan Lake (about 40,000 acres). The Army Corps
of Engineers own the lands under and around Jordan Lake, but state agencies manage most of the
upland areas. Notable state-owned lands within Cape Fear River basin include: a number of
State Parks (Raven Rock, Bay Tree Lake, Bushy Lake State Natural Area, Bald Head Island
State Natural Area, Weymouth Woods, Singletary Lake, Carolina Beach); Wildlife Resources
Commission Game Lands (Holly Shelter Angola Bay, Cape Fear River Wetlands, Suggs Mill
Pond); the Department of Agriculture’s Boiling Springs Lakes Preserve (a joint project with The
Nature Conservancy); the North Carolina Zoo; and the Division of Forest Resources’ Bladen
Lakes State Forest. Also, the Department of Transportation has acquired several properties in
the basin to mitigate for wetlands impacted during highway construction. This property is
permanently protected, and hopefully will be restored to provide a number of ecological benefits.
Local efforts to protect land within the Cape Fear River basin have yielded a great deal of benefit
to water quality. For instance, the City of Fayetteville has protected land along Cross Creek –
land that is an important natural area, a tributary of the Cape Fear, and part of the water supply
for Fayetteville. Also, the North Carolina Botanical Garden Foundation has dedicated as State
Nature Preserves some of the land they own near Chapel Hill, protecting natural diversity and the
water quality of Morgan Creek.
While federal, state and local agencies are important players in land protection efforts within the
Cape Fear River basin, key partners in these efforts are private conservation organizations such
as the North Carolina Coastal Land Trust, Triangle Land Conservancy, Sandhills Area Land
Trust, and the North Carolina Chapter of The Nature Conservancy. Although only partially
shown on the map, these organizations have achieved significant protection in the Cape Fear
River basin. Using innovative tools such as conservation easements, these organizations work
with landowners in a number of ways to protect important natural areas and water quality, as
well as the “open space” of agricultural lands. The work that they do is helping to improve the
quality of life for residents of the Cape Fear River basin.
Chapter 33 – Significant Ecological Resources and Endangered Species 305